Raghu, S., M.L. Smith, and A.M. Simons.  Avoiding dead ends: the experimental evolution of constraint as adaptation in Saccharomyces cerevisiae (Submitted).


Coffey, M.L. and A.M. Simons. 2023. Latitudinal trends in mating system traits in the highly self-fertilizing Lobelia inflata revealed by community science. Ecology & Evolution

Campeau, W., A.M. Simons and B. Stevens. 2023. Intermittent search, not strict Lévy flight, evolves under relaxed foraging distribution constraints. The American Naturalist. DOI: [PDF]

Campeau, W., A.M. Simons and B. Stevens. 2022. The evolutionary maintenance of Lévy flight foraging. PLOS Computational Biology 18(1): e1009490. [PDF]

Morris, R.S., M.E. Compton, and A.M. Simons. 2020. Birth order as a source of within-genotype diversification in the clonal duckweed, Spirodela polyrhiza (Araceae: Lemnoideae). Biological Journal of the Linnean Society 131:1002–1010.

Hitsman, H.W. and A.M. Simons. 2020. Latitudinal variation in norms of reaction of phenology in the greater duckweed Spirodela polyrhizaJournal of Evolutionary Biology 33:1405-1416. DOI: 10.1111/jeb.13678

Wen, L. and A.M. Simons. 2020. Delusions of grandeur: Seed count is not a good fitness proxy under individual variation in phenology. Journal of Evolutionary Biology. 33:1039-1049. DOI: 10.1111/jeb.13633

Côté, K. and A.M. Simons. 2020. Genotype-environment interaction and the maintenance of genetic variation: an empirical study of Lobelia inflata (Campanulaceae). Roy. Soc. Open Sci. 7:191720.

Brady, S. P., …, A.M. Simons et al. 2019. Understanding maladaptation by uniting ecological and evolutionary perspectives. The American Naturalist 194:495-515.

Van Wieren, J.F., and A.M. Simons. 2019. Prescribed fire increases seedling recruitment in a natural pitch pine (Pinus rigida) population at its northern range limit. Natural Areas Journal 39:308-318.

Brady, S. P., …, A.M. Simons et al. 2019. Causes of maladaptation. Evolutionary Applications 12:1229-1242. DOI: 10.1111/eva.12844

Mejbel, H.S., and A.M. Simons. 2018. Aberrant clones: birth order generates life-history diversity in Greater Duckweed, Spirodela polyrhizaEcology and Evolution 8:2021-2031.

Fukase, J., and A.M. Simons. 2016. Increased pollinator activity in urban gardens with more native flora. Applied Ecology and Environmental Research 14:297-310.  DOI: [PDF] The definitive version is available at: {Link}

Hughes, P.W.D., and A.M. Simons. 2015. Microsatellite evidence for obligate autogamy, but abundant genetic variation in the herbaceous monocarp Lobelia inflata. Journal of Evolutionary Biology 28:2068-2077. [PDFThe definitive version is available at: [Link]

Hughes, P.W.D., and A.M. Simons. 2014. Changing reproductive effort within a semelparous reproductive episode. American Journal of Botany 101:1323-1331. DOI:10.3732/ajb.1400283. [Link]

Graham, J.K., M.L. Smith, and A.M. Simons. 2014. Experimental evolution of bet hedging under manipulated environmental uncertainty in Neurospora crassa. Proceedings of the Royal Society B 281: 20140706. [Link]

Simons, A.M. 2014. Playing smart vs. playing safe: the joint expression of phenotypic plasticity and potential bet hedging across and within thermal environments Journal of Evolutionary Biology 27:1047-1056. DOI: 10.1111/jeb.12378 [PDFThe definitive version is available at: [Link]

Hughes, P.W.D., and A.M. Simons. 2014. Secondary reproduction in the herbaceous monocarp Lobelia inflata: time-constrained primary reproduction does not result in increased deferral of reproductive effort. BMC Ecology 14:15. [Link].

Hughes, P.W.D., and A.M. Simons. 2014. The continuum between semelparity and iteroparity: Plastic expression of parity in response to season length manipulation in Lobelia inflataBMC Evolutionary Biology 14:90. [Link].

Hughes, P.W, A.F. Jaworski, C.S. Davis, S.M. Aitken and A.M. Simons. 2014. Development of polymorphic microsatellite markers for Indian Tobacco Lobelia inflata (Campanulaceae)  Applications in Plant Sciences 2: 1300096. [Link].

Simons, A.M. 2011. Modes of response to environmental change and the elusive empirical evidence for bet hedging. Proceedings of the Royal Society B 278:1601-1609. [Link] [PDF]

Simons, A.M., J.M. Goulet, and K.F. Bellehumeur. 2010. The effect of snow depth on overwinter survival in Lobelia inflataOikos 119:1685-1689. [Link] [PDF].

Simons, A.M. 2009. Fluctuating natural selection accounts for the evolution of diversification bet hedging. Proceedings of the Royal Society B 276:1987-1992. [Link]

Wagner, I. and A.M. Simons. 2009. Divergence among arctic and alpine populations of the annual, Koenigia islandica: morphology, life history and phenology. Ecography 32:114-122. [PDF]

Wagner, I. and A.M. Simons. 2009. Divergent norms of reaction to temperature in germination characteristics among populations of the arctic-alpine annual, Koenigia islandicaArctic, Antarctic and Alpine Research 41:388-395.

Wagner, I. and A.M. Simons. 2009. Divergence in germination traits among arctic and alpine populations of Koenigia islandica: light requirements. Plant Ecology 204:145-153.

Simons, A.M. 2008. One big, and many small reasons that direct selection on offspring number is still open for discussion. Journal of Evolutionary biology 21:642-645.

Wagner, I. and A.M. Simons. 2008. Intraspecific divergence in seed germination traits between high and low latitude populations of the arctic-alpine annual Koenigia islandicaArctic, Antarctic and Alpine Research 40:233-239.

Simons, A.M. 2007. Selection for increased allocation to offspring number under environmental unpredictability. Journal of Evolutionary biology 20:813-817. [PDFThe definitive version is available at: [Link]

Simons, A.M., and I. Wagner. 2007. The characterization of complex continuous norms of reaction. Oikos 116:986-994.

Simons, A.M., and M.O. Johnston 2006. Environmental and genetic sources of diversification in the timing of seed germination: Implications for the evolution of bet hedging. Evolution 60:2280-2292.

Simons, A.M. 2004. Many wrongs: the advantage of group navigation. Trends in Ecology and Evolution 19:453-455.

Simons, A.M. 2003. Invasive aliens and sampling bias. Ecology Letters 6:278-280.

Simons, A.M., and M.O. Johnston 2003. Suboptimal timing of reproduction in Lobelia inflata may be a conservative bet-hedging strategy. Journal of Evolutionary Biology 16:233-243. [PDFThe definitive version is available at: [Link]

Simons, A.M. 2002. The continuity of microevolution and macroevolution. Journal of Evolutionary Biology 15:688-701. [PDFThe definitive version is available at: [Link]

Simons, A.M., and M.O. Johnston. 2000. Plasticity and the genetics of reproductive behaviour in the monocarpic perennial, Lobelia inflata (Indian tobacco). Heredity 85:356-365.

Simons, A.M., and M.O. Johnston. 2000. Variation in seed traits of Lobelia inflata (Campanulaceae): sources and fitness consequences. American Journal of Botany 87:124-132.

Simons, A.M., and M.O. Johnston. 1999. The cost of compensation. American Naturalist 153:683-687.

Simons, A.M., Y. Carrière, and D.A. Roff. 1998. The quantitative genetics of growth in a field cricket. Journal of Evolutionary Biology 11:721-734.

Simons, A.M., and M.O. Johnston. 1997. Developmental instability as a bet-hedging strategy. Oikos 80:401-406. [PDF] [Link].

Roff, D.A., and A.M. Simons. 1997. The quantitative genetics of wing dimorphism under laboratory and ‘field’ conditions in the cricket Gryllus pennsylvanicusHeredity 78:235-240.

Carrière, Y., A.M. Simons, and D.A. Roff. 1996. The effect of the timing of post-diapause egg development on survival, growth, and body size in Gryllus pennsylvanicusOikos 75:463-470.

Simons, A.M., and D.A. Roff. 1996. The effect of a variable environment on the genetic correlation structure in a field cricket. Evolution 50:267-275.

Gasol, J.M., A.M. Simons, and J. Kalff. 1995. Patterns in the top-down versus bottom-up regulation of heterotrophic nanoflagellates in temperate lakes. Journal of Plankton Research 17:1879-1903.

Simons, A.M., and D.A. Roff. 1994. The effect of environmental variability on the heritabilities of traits of a field cricket. Evolution 48:1637-1649.